Mixed broadleaved forests typical of less-frequently flooded, well-aerated mineral soils on floodplains of the middle and lower reaches of major European rivers. The canopy in high-forest stands can be very tall and multi-layered and is typically dominated by various mixtures of Alnus glutinosa, Fraxinus angustifolia, Fraxinus excelsior, Populus alba, Populus canescens, Prunus padus, Quercus robur, Ulmus glabra, Ulmus laevis and Ulmus minor. There is typically an abundant and varied understorey, again often structurally complex, with a range of small trees, shrubs and lianas that are more typical of mesic deciduous forests (such as T1E Carpinus and Quercus mesic deciduous forest).