Hemiboreal pine and birch-pine herb-rich open forests on fertile soils of the Southern Urals and Southern Siberia, and relict birch-poplar forests of Europe
Vascular plants: Achillea millefolium subsp. millefolium, Aconitum lycoctonum subsp. lycoctonum, Actaea europaea, Adenophora liliifolia, Agrimonia pilosa, Aster alpinus, Betula pendula var. pendula, Brachypodium pinnatum, Bupleurum longifolium, Calamagrostis arundinacea, Campanula glomerata, Caragana frutex, Carex digitata, Carex pediformis subsp. rhizodes, Cephalaria uralensis, Cerastium pauciflorum, Conioselinum tataricum, Cotoneaster laxiflorus, Cytisus ruthenicus, Digitalis grandiflora, Dracocephalum ruyschiana, Euphorbia subcordata, Filipendula vulgaris, Fragaria vesca, Galium boreale, Geranium pseudosibiricum, Gypsophila altissima, Hieracium umbellatum, Larix sibirica, Lathyrus gmelinii, Lathyrus pisiformis, Lathyrus vernus, Lilium martagon var. pilosiusculum, Luzula pilosa, Melica nutans, Milium effusum subsp. effusum, Origanum vulgare subsp. vulgare, Phlomis tuberosa, Pinus sylvestris, Pleurospermum uralense, Poa nemoralis subsp. nemoralis, Polygonatum odoratum, Primula veris subsp. macrocalyx, Prunus fruticosa, Psephellus sibiricus, Pulmonaria mollis subsp. mollis, Pulsatilla patens, Rosa majalis, Rubus saxatilis, Sanguisorba officinalis, Serratula coronata, Seseli libanotis, Silene chalcedonica, Silene nutans subsp. nutans, Solidago virgaurea subsp. virgaurea, Sorbus aucuparia subsp. aucuparia, Stachys officinalis subsp. officinalis, Succisa pratensis, Thalictrum foetidum, Thalictrum minus subsp. minus, Trientalis europaea, Trifolium lupinaster, Trifolium medium subsp. medium, Trollius europaeus, Veronica chamaedrys subsp. chamaedrys, Vicia sepium, Viola canina, Viola collina, Viola mirabilis, Viola rupestris